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The large citrus fruit of today evolved originally from small, edible berries over millions of years.Citrus plants diverged from a common ancestor about 15 million years ago, which was about when it diverged from the closely related severinia, for example the Chinese box orange.From the endocarp, string-like "hairs" extend into the locules, which provide nourishment to the fruit as it develops.Citrus fruits are notable for their fragrance, partly due to flavonoids and limonoids (which in turn are terpenes) contained in the rind, and most are juice-laden.Something similar has occurred with the wide array of chili peppers originating in the hybridization of a few initial species.Hybrids are expected to plot between their parents.The outermost layer of the pericarp is an "exocarp" called the flavedo, commonly referred to as the zest.The middle layer of the pericarp is the mesocarp, which in citrus fruits consists of the white, spongy "albedo", or "pith".

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The generic name originated from Latin, where it referred to either the plant now known as Citron (C. It is somehow related to the ancient Greek word for cedar, κέδρος (kédros).Fruits produced with organic agriculture have been shown to contain more vitamin C than those produced with conventional agriculture in the Algarve, but results depended on the species and cultivar.

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